arXiv2026
Many natural ecosystems harbor large numbers of coexisting species competing for far fewer distinct resources, in apparent defiance of the competitive exclusion principle. Various mechanisms have been proposed to explain this apparent paradox, often pertaining to organisms with a two-stage sessile--propagule life cycle. Here we develop a stochastic model class for such propagule disperser communities that combines competition--colonization trade-offs, spatial heterogeneity, demographic stochasticity, as well as inherited trait variation, and recover several classical models as special or limiting cases. Using bifurcation analysis, we classify equilibrium coalitions near the extinction threshold and give sufficient conditions for their realization by macroscopic equilibria away from the threshold, bypassing the costly numerical computation of the actual equilibrium states. Illustrative examples examine the resulting trait distributions and coalition patterns, demonstrating the interactive effects of different coexistence mechanisms.